Rhexoxylon cortaderitaense (Menéndez) comb. nov., a species of permineralized stems newly assigned to the Corystospermaceae, from the Triassic of Argentina

Abstract

Anatomically preserved stems of Late Triassic corystosperms from the upper Cortaderita Formation in San Juan province, Argentina are described and assigned to Rhexoxylon cortaderitaense (Mene´ndez) comb. nov. These specimens were originally attributed to conifers, a group not closely related to corystosperms. The silicified axes preserve features of the cortex, pith, and secondary vascular system. Like all Rhexoxylon species, these axes have two discontinuous cambial rings that develop centrifugal secondary xylem (normal secondary xylem) and centripetal secondary xylem (inverted secondary xylem). Both types of xylem are separated by a band of parenchymatous tissue developed by a remnant cambium. The centrifugal xylem is divided by parenchymatous rays following differential cambial activity. The secondary xylem is pycnoxylic with uniseriate rays; tracheids have mainly biseriate and alternate bordered pits on radial and tangential walls, and one simple oval pit in cross-fields. The distinctive feature of R. cortaderitaense is the conspicuous amount of centripetal secondary vascular tissues not forming perimedullar bundles, this being interpreted as an apomorphic trait. This interpretation of the character modifies previous hypothesized phylogenetic relationships of corystosperm wood taxa.